|
Kernel Set Scuttlebutt - (Bob Nielsen)
“Scuttlebutt”: The cask of drinking water on ships was called a scuttlebutt and since sailors exchanged gossip when they gathered at the scuttlebutt for a drink of water, scuttlebutt became U.S. Navy slang for gossip or rumors. A butt was a wooden cask, which held water or other liquids; to scuttle is to drill a hole, as for tapping a cask.
Nautical Terms and Phrases, NAVAL HISTORICAL CENTER, Washington DC 20374-5060.
Online at www.ussbrainedd630.com/terms.htm [URL verified 8/8/05].
The scuttlebutt heard in many coffee shops in Indiana these days is that folks are walking corn fields and finding a gamut of kernel set success ranging from excellent to pathetic. Growers’ interest in this topic obviously lies with the fact that the number of kernels per ear is a rather important component of total grain yield per acre for corn.
Poor kernel set, meaning unacceptably low kernel numbers, is a bad omen for fields that otherwise appear to be in good shape. Good or poor kernel set is determined from pollination through the early stages of kernel development; typically 2 to 3 weeks post-pollination.
Problems with kernel set stem from ineffective pollination, kernel abortion, or both. Distinguishing between these two symptoms is easy. Determining the exact cause of the problem is often difficult.
Potential Yield Loss
The potential loss in grain yield caused by lower kernel numbers per ear can be estimated using the formula of the so-called Yield Component Method first described by the Univ. of Illinois many years ago (Univ. of Illinois, 2005). For example, the loss of only 1 kernel per row for a hybrid with 16-row ears and a stand count of 30,000 ears per acre would equal a potential yield loss of approximately 5 bushels per acre (1 [kernel] x 16 [rows] x 30 [thousand ears per acre] divided by 90 [thousand kernels per bushel]).
Ineffective Pollination
Poor kernel set may be caused by ineffective pollination and the subsequent failure to fertilize ovules on the cob. Ineffective pollination is characterized by an absence of noticeable kernel development. In other words, all you see is cob tissue. Pollination problems may be due to several stress factors this year, sometimes working together to influence kernel set.
Severe drought stress, aggravated by excessive heat, can delay silk emergence to the extent that pollen shed is complete or nearly complete by the time the silks finally emerge from the husk. Without a pollen source, ovule fertilization cannot occur.
Persistent severe silk clipping by insects such as the corn rootworm beetle or Japanese beetle throughout the active pollen shed period can also limit the success of pollination. The simultaneous effects of severe drought stress on silk emergence can easily amplify the consequences of severe silk clipping.
Severe drought stress coupled with excessive heat and low humidity can sometimes desiccate emerged silks to the point that they are no longer receptive to pollen grain germination. I suspect this is low on the list of possible stressors for Indiana most years (because of our typically high humidity levels), but may have played a role in some fields this year. Similarly, I doubt that pollen viability was an issue for Indiana cornfields because temperatures in the low 90’s are usually not great enough to kill pollen.
Consecutive days of persistent rainfall or showers that keep tassels wet for many hours of a day can delay or interfere with anther exsertion and pollen shed. Remember the remnants of Hurricane Dennis that visited many parts of Indiana earlier last month? I admit to not having a lot of evidence to support the following conjecture, but I suspect that some of the simply weird patterns of poor kernel set evident in some fields that were trying to pollinate during that week may have been related to those many days of showery humid weather (coupled with the excessive cloudiness and its effect on photosynthesis).
Exceptionally long potential ears resulting from good weather during ear size determination (not an issue for some Indiana fields this year) sometimes fail to pollinate the final kernels near the tip of the cob. Remember, butt silks emerge first and tip silks emerge last. With oversized ears, sometimes those tip silks emerge after all the pollen has been shed. See the “Word to the Wise” below.
Kernel Abortion
Poor kernel set can also be a reflection of kernel abortion following successful fertilization of the ovules on the cob. In contrast to ineffective pollination, initial kernel development obviously precedes kernel abortion, so the symptoms are usually shriveled remnants of kernels that may be whitish- or yellowish-translucent.
Kernel abortion results from severe stresses that greatly reduce the overall photosynthetic output of the plant very early in the post-pollination grain-filling period. Obvious photosynthetic stressors include severe drought & heat stress, consecutive days of excessively cloudy weather and significant loss of photosynthetically active leaf area (e.g., hail damage, leaf diseases, insect damage, nutrient deficiency).
Warm nights during pollination and early grain fill may indirectly affect survival of developing kernels. Research suggests that the increased rate of kernel development due to warmer temperatures lowers the available amount of photosynthate per unit of thermal time; which then becomes a stressor to kernel development particularly at the tip of the ear, leading to kernel abortion (Cantarero et. al., 1999).
Final Word to the Wise
A plethora (meaning a whole lot) of blank cob tips can quickly ruin the joy of walking a cornfield in the middle of August. Before getting too bent out of shape over the missing kernels, remember to count the number of harvestable kernels on those ears. I’ve been amazed at the number of fields I’ve walked the past week whose ears exhibit 1 to 2 inches of blank tips, yet still contain 16 rows by 30 to 35 harvestable kernels per row. Those are perfectly acceptable ear sizes in a year where dry weather has been a concern.
Related References
Cantarero, M.G., A.G. Cirilo, and F.H. Andrade. 1999. Night temperature at silking affects kernel set in maize. Crop Sci 39:703-710.
Nielsen, R.L. (Bob). 2004a. Estimating Corn Grain Yield Prior to Harvest. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.04/YieldEst-0718.html.[URL verified 8/8/05].
Nielsen, R.L. (Bob). 2004b. Grain Fill Stages in Corn. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.04/GrainFill-0705.html. [URL verified 8/8/05].
Nielsen, R.L. (Bob). 2004c. Yield Loss Potential During Grain Fill. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.04/GrainFillStress-0705.html. [URL verified 8/8/05].
Nielsen, R.L. (Bob). 2005b. Silk Emergence. Corny News Network, Purdue Univ. Available online at www.kingcorn.org/news/articles.05/Silks-0704.html. [URL verified 7/4/05].
Nielsen, R.L. (Bob). 2005c. Tassel Emergence & Pollen Shed. Corny News Network, Purdue Univ. Available online at www.kingcorn.org/news/articles.05/Tassels-0704.html.[URL verified 7/4/05].
Univ. of Illinois. 2005. Estimating Corn Yields. (An Online Calculator). Illinois Agronomy Handbook. Online at www.ag.uiuc.edu/iah/index.php?ch=ch2/est_corn_yield.html. [URL verified 8/8/05].
back to top

Estimating Corn Grain Yield Prior to Harvest - (Bob Nielsen)
Fancy colored yield maps are fine for verifying grain yields at the end of the harvest season, but bragging rights for the highest corn yields are established earlier than that down at the Main Street Cafe, on the corner of 5th and Earl. Corn in IndianaSome patrons of the cafe begin “eyeballing” their yields as soon as their crops reach “roasting ear” stage. Some of the guys there are pretty good (or just plain lucky) at estimating yields prior to harvest, while the estimates by others are not even close to being within the proverbial ballpark. Interestingly, they all use the same procedure referred to as the Yield Component Method.
Other pre-harvest yield prediction methods exist (Lauer, 2002; Thomison, 2005), but the Yield Component Method is probably the most popular because it can be used well ahead of harvest; as early as the so-called “roasting ear” or milk (R3) stage of kernel development. Under “normal” conditions, the kernel milk stage occurs about 18 to 22 days after pollination is complete (Nielsen, 2004a). Estimates made earlier in the kernel development period risk being overly optimistic if subsequent severe stresses cause unforeseen kernel abortion (Nielsen, 2004b).
The Yield Component Method was originally described by the University of Illinois many years ago (Univ. of Illinois, 2005) and is based on the premise that one can estimate grain yield from estimates of the yield components that constitute grain yield. These yield components include number of ears per acre, number of kernel rows per ear, number of kernels per row, and weight per kernel. The first three yield components (ear number, kernel rows, kernels/row) are easily measured in the field.
Final weight per kernel obviously cannot be measured until the grain is mature (kernel black layer) and, realistically, at harvest moisture. Consequently, an average value for kernel weight, expressed as 90,000 kernels per 56 lb bushel, is used as a proverbial “fudge factor” in the yield estimation equation.
Crop uniformity greatly influences the accuracy of any yield estimation technique. The less uniform the field, the greater the number of samples that should be taken to estimate yield for the field. There is a fine line between fairly sampling disparate areas of the field and sampling randomly within a field so as not to unfairly bias the yield estimates up or down.
1. At each estimation site, measure off a length of row equal to 1/1000th acre. For 30-inch (2.5 feet) rows, this equals 17.4 feet.
TIP: For other row spacings, divide 43,560 by the row spacing (in feet) and then divide that result by 1000 (e.g., [43,560/2.5]/1000 = 17.4 ft).
2. Count and record the number of ears on the plants in the 1/1000th acre of row that you deem to be harvestable.
TIP: Do not count dropped ears or those on severely lodged plants unless you are confident that the combine header will be able to retrieve them.
3. For every fifth ear in the sample row, record the number of complete kernel rows per ear and average number of kernels per row. Then multiply each ear’s row number by its number of kernels per row to calculate the total number of kernels for each ear.
TIPS: Do not sample nubbins or obviously odd ears, unless they fairly represent the sample area. If row number changes from butt to tip (e.g., pinched ears due to stress), estimate an average row number for the ear. Don’t count the extreme butt or tip kernels, but rather begin and end where you perceive there are complete “rings” of kernels around the cob. Do not count aborted kernels. If kernel numbers are uneven among the rows of an ear, estimate an average value for kernel number per row.
4. Calculate the average number of kernels per ear by summing the values for all the sampled ears and dividing by the number of ears.
EXAMPLE: For five sample ears with 480, 500, 450, 600, and 525 kernels per ear, the average number of kernels per ear would be (480 + 500 + 450 + 600 + 525) divided by 5 = 511.
5. Estimate the yield for each site by multiplying the ear number by the average number of kernels per ear, then dividing that result by 90. The value of ‘90’ represents the average number of kernels (90,000) in a bushel of corn.
TIP: Use a lower value (e.g., 80) if grain fill conditions have been excellent (larger kernels, fewer per bushel) or a larger value (e.g., 100) if grain fill conditions have been stressful (smaller kernels, more per bushel).
Example
Let’s say you counted 30 harvestable ears at the first sampling site. Let’s also assume that the average number of kernels per ear, based on sampling every 5th ear in the sampling row, was 511. The estimated yield for that site would (30 x 511) divided by 90, which equals 170 bu./ac.
Repeat the procedure throughout field as many times as you deem to be representative. Calculate the average yield for all the sites to estimate the yield for the field.
Remember that this method for estimating pre-harvest grain yield in corn indeed provides only an estimate. Since kernel size and weight will vary depending on hybrid and environment, this yield estimator should only be used to determine “ballpark” grain yields. Yield will be overestimated in a year with poor grain fill conditions (e.g., low kernel size and weight from a drought year) and underestimated in a year with excellent grain fill conditions (e.g., larger kernel size and weight from non-stress grain fill periods).
You can try to improve the yield estimation for unusual grain fill conditions by adjusting the estimation formula . For example, if you believe that kernel weight will be lower due to stress during grain fill, you may elect to replace the value of “90” in the equation with “100” to reflect the potential for smaller and lighter kernels (i.e., more kernels per 56 lb. bushel). Conversely, in a good crop year, you may elect to replace the value of “90” in the equation with “80” to reflect the potential for larger and heavier kernels (i.e., fewer kernels per 56 lb. bushel).
Recognize that the Yield Component Method for estimating corn grain yield is probably only accurate within plus or minus 30 bushels of the actual yield. Obviously, the more samples you measure within a field, the more accurately you will “capture” the variability of yield throughout the field. Use the yield estimates obtained by this method for general planning purposes only.
Related References
Lauer, Joe. 2002. Methods for Calculating Corn Yield. Agronomy Advice, Univ. of Wisconsin-Madison. Online at http://corn.agronomy.wisc.edu/AAdvice/2002/A033.pdf. [URL verified 8/9/05].
Nielsen, R.L. (Bob). 2004a. Grain Fill Stages in Corn. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.04/GrainFill-0705.html. [URL verified 8/9/05].
Nielsen, R.L. (Bob). 2004b. Yield Loss Potential During Grain Fill. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.04/GrainFillStress-0705.html. [URL verified 8/9/05].
Nielsen, R.L. (Bob). 2005. Kernel Set Scuttlebutt. Corny News Network, Purdue Univ. Online at www.kingcorn.org/news/articles.05/KernelSet-0809.html. [URL verified 8/9/05].
Thomison, Peter. 2005. Estimating Yield Losses in Drought Damaged Corn Fields. C.O.R.N. Newsletter, Ohio State Univ. Online at http://corn.osu.edu/story.php?setissueID=96&storyID=571 [URL verified 8/9/05].
Univ. of Illinois. 2005. Estimating Corn Yields. (An Online Calculator). Illinois Agronomy Handbook. Online at www.ag.uiuc.edu/iah/index.php?ch=ch2/est_corn_yield.html. [URL verified 8/9/05].
For other Corny News Network articles, browse through the CNN Archives at www.kingcorn.org/news/archive.html. For other information about corn, take a look at the Corn Growers’ Guidebook at www.kingcorn.org.
back to top

Impact of Warm Night Temperatures on Corn Grain Yields – (Peter Thomison, Ohio State University)
When we experience a hot, dry growing season like that of 2005, we often focus our attention on the appearance of stressed plants (leaf rolling, leaf firing, poor canopy closure), and relate stress symptoms or injury to lower yield potential. However warm night temperatures, which are often associated with drought, also adversely affect yield potential. High night temperatures (in the 70s or 80s) result in wasteful respiration and a lower amount of dry matter accumulation in plants. With high night temperatures more of the sugars produced by photosynthesis during the day are lost; less is available to fill developing kernels grain, thereby lowering potential grain yield. Past research indicates that corn grown at night temperatures in the mid 60s outyields corn grown at temperatures in the mid 80s. Corn yields are often higher with irrigation in western states, which have low humidity and limited rainfall. While these areas are characterized by hot sunny days, night temperatures are often cooler than in the Eastern Corn Belt. Low night temperatures account in part for our record high corn yields in 2004. During most of the 2004 growing season, temperatures were below normal. From late June through most of August, a period of time that included most of the grain fill period, weekly temperatures were cooler than normal - as much as 4 to 7 degrees below normal in August. Cool night temperatures in 2004 reduced respiration losses during grain fill. The absence of moisture stress was especially important during grain filling. In parts of the Ohio where rainfall was below average during grain fill in July and August, cooler than average temperatures minimized moisture stress. The high corn yields of 2003 were also associated with cooler than normal night temperatures during the grain fill period.
Impact of drought on grain composition
Drier and warmer than normal conditions during late vegetative development and grain fill also influence grain composition. When drought stress conditions severely depress corn yields, protein deposited early in the kernels is less diluted by starch deposited later during grain fill; consequently grain protein concentration increases. Conversely, optimum soil moisture, whether from rain or irrigation, promotes filling of kernels with starch and reduces protein content.
back to top

Assessing Potential For Nitrate Problems in Corn – (Peter Thomison, Ohio State University)
Hot dry weather across Ohio has raised questions concerning the potential for toxic levels of nitrates in corn harvested for silage. Nitrates absorbed from the soil by plant roots are normally incorporated into plant tissue as amino acids, proteins and other nitrogenous compounds. Thus, the concentration of nitrate in the plant is usually low. The primary site for converting nitrates to these products is in growing green leaves. Under unfavorable growing conditions, especially drought, this conversion process is retarded, causing nitrate to accumulate in the stalks, stems and other conductive tissue. The highest concentration of nitrates is in the lower part of the stalk or stem. For example, the bulk of the nitrate in a drought-stricken corn plant can be found in the bottom third of the stalk. If moisture conditions improve, the conversion process accelerates and within a few days nitrate levels in the plant returns too normal.
The highest levels of nitrate accumulate when drought occurs during a period of heavy nitrate uptake by the corn plant. A drought during or immediately after pollination is often associated with the highest accumulations of nitrates. Extended drought prior to pollination is not necessarily a prelude to high accumulations of nitrate. The resumption of normal plant growth from a heavy rainfall will reduce nitrate accumulation in corn plants, and harvest should be delayed for at least 1 to 2 weeks after the rainfall. Not all drought conditions cause high nitrate levels in plant. If the supply of soil nitrates is in the dry soil surface, plant roots will not absorb nitrates. Some soil moisture is necessary for absorption and accumulation of the nitrates.
If growers want to salvage part of their drought damaged corn crop as silage, it’s best to delay harvesting to maximize grain filling, if ears have formed. Even though leaves may be dying, the stalk and ear often have enough extra water for good fill. Kernels will continue to fill and the increases in dry matter will more than compensate for leaf loss unless plants are actually dying or dead. Moreover if nitrate levels are high or questionable, they will decrease as plant get older and nitrates are converted to proteins in the ear.
For information on testing and feeding corn with varying nitrate-nitrogen levels, check out the following:
Drought-Stressed Corn For Silage - Bill Weiss available on-line at http://corn.osu.edu/archive/2002/jul/02-24.html#linka.
Nitrates in Dairy Rations - Maurice Eastridge and Bill Weiss, Ohio State University Fact Sheet AS-0003-99 available on-line at http://ohioline.ag.ohio-state.edu/as-fact/0003.html.
|